Ph.D. Biología, Graduate School of Arts and Sciences, Harvard University
M.Sc. Biología, Graduate School of Arts and Sciences, Harvard University
B.Sc. Biología, Universidad de los Andes
His research interest is on the origin and preservation of biodiversity in a systematic evolutionary context. The basis of his research are studies on flora and fauna, plant ecology, animals and their interaction, mainly but not exclusively on the Neotropical region.
Through his work, Professor Madriñan formulates hypothesis on kingship pattern of different taxonomic levels organism, framed in systematic phylogenetics. His work is a contribution to different biology fields (e.g. sistematic, paleontology, ecology, ethology, morphology, embriology, fisiology, molecular biology, genetics, etc.). His research also contributes to the study of the history and causes of evolution. He applies biota´s systematic studies to biogeographic history, conservation biology and studies on biodiversity.